Earthworms: the most important soil and ecosystem engineers
Earthworms are thought to be the most ancient soil animals having started colonizing terrestrial environments about 600 million years ago (Spain and Lavelle 2001). They are the most predominant soil fauna except in dry and cold climates. Earthworms are semiaquatic animals which extract water continuously from the surrounding soil inorder to maintain their cuticle in a moist state to facilitate gas exchange. Thus moisture status is a major limitation to earthworm activities and distribution. Spain and Lavelle (2001) reported that since earthworms live in direct and continuous contact with the soil matrix and the soil solution, their presistence, propagation and activity are greatly affected by the chemical (pH, dissolved ions) characteristics of the soil. Based on their sensitivity to soil pH, earthworms are grouped into acidophilic species (able to thrive below pH 6 such as in organic forest litter), neutrophilic species (they prefer soil pH 6 to 7) and basophi lic species (prefer b...